Le Vell is charged under his real name of Michael Turner with five counts of rape, three of indecent assault, two of sexual activity with a child and two of causing a child to engage in sexual activity.
The 48-year-old, best known as the soap's Kevin Webster, denies all 12 charges against him.
The alleged victim, who cannot be identified, was cross-examined by Le Vell's defence team on Tuesday over alleged inconsistencies in her statements to police and accused of telling "a little lie".
Her mum later wept in the witness box as she recalled being told by her daughter about the allegations for the first time.
"I believe he (Le Vell) had molested and abused her on a regular basis and there have been a number of rapes," she said.
Describing how she went to Le Vell over the claims, she told the court that she had said to him: "She's made an accusation. You raped her when she was younger.
"Before I finished that sentence, he said: 'Oh, you're joking! This is a joke! It's a joke! You're having a laugh!'"
Prosecution lawyer Eleanor Laws QC asked the witness: "Did he ask what she said?"
The girl's mother replied: "He never asked what she said."
She told the jury that her daughter had described how Le Vell would cover her face with a teddy bear while sexually abusing her, adding: "In the car, she said: 'It's happened a lot more than I'm making out, mummy. It's a lot worse.' She said: 'Mummy, you don't want to know.'"
Le Vell's defence lawyer Alisdair Williamson asked the witness about a call she made to police, asking officers if she could sell her story to the papers.
The witness replied: "I chose not to do that."
During his earlier cross-examination of the girl at the centre of the case, Mr Williamson had told the court that she had initially told officers that she had never talked to anyone else about the alleged sexual abuse - when she had told some friends.
I told two friends because they saw me crying ...
Mr Williamson further suggested that, when the girl was interviewed on video by police, she had said some "silly things" that she had left out of her account given in court.
He told the court that she had claimed to remember the exact times when alleged rapes took place, but that the details meant two of the attacks would have happened at the exact same time.
"Was it a detail you were putting in to make you more convincing and you now realise it sounds a bit silly?" the lawyer asked.
"No," she replied.
"It didn't happen," Mr Williamson said.
"It did," the witness replied.
The lawyer told the court that the witness had been to an "inspirational self-help" conference and listened to a woman's account of being raped at a young age and then rebuilding her life.
"Is that what some of this is about?" the lawyer asked.
"You heard this lady talking about how she was raped when she was very young and how she went on and became a model? You want to become an actress or a dancer, is that right?"
She replied: "I wanted to - I don't anymore."
The jury heard that the girl had made further allegations against Le Vell in the months after the Crown Prosecution Service decided to drop the case.
When questioned why that was, she broke down in tears and said: "I had a lot more flashbacks. I was very confused because what he did to me was so wrong. I was so young."
Prosecution lawyer Eleanor Laws QC asked: "Has Michael Le Vell done anything to cause you to make up these wicked lies?"
"No," the girl said.
"Why have you made these allegations?" Ms Laws asked.
"Because they are true," she replied.
Concluding the girl's evidence in the witness box, Ms Laws asked: "Have you some sort of vendetta, any reason to hate Michael Le Vell or bring him down and cause him all of this?"
"No," the girl replied.
The trial is set to continue on Wednesday afternoon.